Waardenburg syndrome

Definition
Waardenburg syndrome (WS) encompasses several different
hereditary disorders, the main features of which variably include abnormal
pigmentation, hearing loss, and a subtle difference in facial features.
Certain other physical anomalies occur less frequently in WS.
Description
In 1951, Dr. Petrus Waardenburg reported a syndrome of
dystopia canthorum, heterochromia of the irides, and hearing loss.
Dystopia canthorum (also called telecanthus) describes a subtle but
unusual facial feature in which the inner corners of the eyes (canthi) are
spaced farther apart than normal, yet the eyes (pupils) themselves are
normally spaced. The result is that the eyes appear to be widely spaced,
even though they are not. Heterochromia means different-colored, and
irides is the plural form of iris-;the colored portion of the eye. Thus,
someone with heterochromia of the irides has differentcolored eyes, often
one brown and one blue. Another feature not originally noted by Dr.
Waardenburg, but now considered a major sign of WS is a white forelock
(white patch of hair extending back from the front of the scalp). In fact,
disturbances in pigmentation (coloring) of various parts of the body are
consistent features of WS. Uncommon but serious physical anomalies
associated with WS include Hirschprung disease (intestinal malformation),
spina bifida, cleft lip/palate, and musculoskeletal abnormalities of the
arms.
Five types of WS have been defined based on clinical
symptoms or genetic linkage. As of 2000, six different genes were
associated with WS. Most families show autosomal dominant inheritance, but
autosomal recessive inheritance and sporadic (single) cases are also seen.
People with WS are not at increased risk for mental retardation, and
vision loss is not more common. For the majority of those with WS, hearing
loss is the only major medical problem they will have.
WS1 is the "classic" form of WS, and if someone uses just
the name Waardenburg syndrome (with no modifying number), they are most
likely referring to the group of disorders as a whole or just WS1. WS2 may
occasionally be referred to as WS without dystopia canthorum. WS3 is also
known as Klein-Waardenburg syndrome, as well as WS with upper limb
anomalies. Alternate names for WS4 include Waardenburg-Hirschprung
disease, Waardenburg-Shah syndrome, Shah-Waardenburg syndrome, and
Hirschprung disease with pigmentary anomaly.
Genetic profile
Since Dr. Waardenburg's original description of his
patients in 1951, many more families with the same or similar symptoms
have been reported. By 1971, it became clear that a proportion of families
have WS without dystopia canthorum. At that point, Waardenburg syndrome
was divided into two distinct types, WS1 and WS2. In addition, a few
individuals with typical signs of WS1 were found to also have
musculoskeletal symptoms. This form of the disorder was named
Klein-Waardenburg syndrome, now also known as WS3. Further, some
researchers noted yet a different pattern of anomalies involving
pigmentation defects and Hirschprung disease, which eventually became
known as WS4. Finally, genetic testing of WS2 families has shown at least
two subtypes-;those that show genetic linkage are designated as WS2A and
WS2B.
The four major types of WS have all been studied through
DNA (genetic) analysis. There is some agreement between the clinical
subtypes of WS and mutations in different genes, but genetic analysis has
also served to confuse the naming scheme somewhat. The different types of
WS, their inheritance patterns, and the genes associated with them, are
listed below.
WS1
A number of different mutations in a single copy of the
PAX3 gene on chromosome 2 are responsible for all cases of WS1, meaning it
is always inherited as an autosomal dominant trait. The PAX3 gene plays a
role in regulating other genes that have some function in producing
melanocytes (pigment-producing cells). PAX3 was formerly known as the HUP2
gene.
WS2A
People who have typical signs of WS2 are designated as
having WS2A only if genetic testing shows them to have a mutation in the
MITF gene on chromosome 3. As with WS1, all cases of WS2A appear to be
autosomal dominant. There is evidence that MITF is one of the genes
regulated by PAX3.
WS2B
Some individuals with typical WS2 have had normal MITF
gene analysis. A search for a different WS2 gene showed that some cases
are linked to a gene on chromosome 1. This gene has been tentatively
designated WS2B until its exact chromosomal location and protein product
are identified. WS2B displays autosomal dominant inheritance.
WS3
Several people with a severe form of WS1 have been shown
by genetic analysis to have a deletion of a small section of chromosome 2.
Several genes are located in this section, including the PAX3 gene. Not
all patients with WS3 have had the exact same genetic anomaly on
chromosome 2, which may explain the variation in symptoms that have been
reported. Some families with WS3 have displayed autosomal dominant
inheritance, while other individuals with the condition have been sporadic
cases.
Source: http://health.families.com/waardenburg-syndrome-1183-1187-gecd